How to access the gold and items your heroic ancestors left behind
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Unfortunately, you can no longer access the chest in the Chamber of Fate, unless you have Fable II: Limited Edition.
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Now with Fable II Limited Edition, if you do have the limited edition you can opened as soon as you reach the Guild. From what I have read on the web, this addition is the only way to open the chest now, though there may be a way to get the items within the chest through downloadable content.