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Spike is from the first Ape Escape game, taking the leading role in that product. He is a fun and diverse character, assisted by his Gotcha gadgets, which include a remote-controlled car! He is also able to call upon monkeys to help in battle. It is important to time Spike's attacks correctly, as if one of the Gotcha gadgets misses, for example, it will leave him defenseless.
PaRappa and Spike are rivals because Spike is concerned that PaRappa is working with the monkeys in trying to take over the world.