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To inflict damage on this type of enemy it is best to use your magical towers as magic damage bypasses armor ensuring that your foe will take a good beating as they march towards the exit point.
Aim your Rain of Fire reinforcement just ahead of the marching enemy as it takes a little while for it to land at it's destination.
Help your soldiers in battle by using your archers to send a rain of arrows into the enemy when you attack.
It is important that these are overlooked by multiple combinations of Mage and Archer Towers.
In key strangle points a good tactic is to double up your barracks. This is done by positioning two militia barracks close to each other and aligning the flag rally points so it doubles up on strength and attack power.
If you find yourself being overwhelmed and your barracks soldiers about to die you can upgrade them mid-battle so they are instantly retrained and tougher with full health.
It is crucial that you keep soldiers away from Parasytes otherwise you are just feeding them so they grow into Reapers.
Reinforcements should never be on Cooldown when there are enemies on the screen as it is important that you keep your current troops alive as long as possible and reinforcements although not particularly tough are always capable of turning the tide of battle to your favour.